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The remaining authors declare that they have no conflict of interests

Clinical manifestations, diagnosis and epidemiological studies of ulcerative colitis UC, first described in 1859, is usually confined to the colon, where persistent superficial inflammation of the upper mucosa of the colon develops, leading to colonic ulceration and bleeding

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These include copy number variants or larger deletions and insertions, noncoding transcripts such as lncRNAs and microRNAs as modulators of gene expression and disease, and mitochondrial gene variants 21,87

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